Eye of the Tiger

Question: Could a single Animal Feature count for two or more abilities, assuming that you pay the requisite Legend and the connection makes sense (example, Eagle eyes adding to both Awareness and Marksmanship, or a wolf nose adding to Investigation and Survival)?

Yes, but also no. Really, it's a Storyteller call that depends entirely on what you're doing and why.

If you have an idea for different Animal Features that are in some way unique, interesting and fun and want them pseudo-combined because it makes for cool flavor, that is totally doable. Maybe one of your eyes adds to Awareness and the other to Marksmanship, and they look slightly different for stunting awesomeness; maybe you want to grow two separate extra rows of teeth to grant you Brawl and Presence because you're terrifying and dangerous. That kind of things's fine, especially as you gain more Animal boons and therefore need to come up with more features, and especially at god-level if you have Bestial Nature and have already filled up some of those "slots" with permanent animal parts.

What's not doable is when features are combined in such a way as to allow the Scion to overall be less animalistic. If you're trying to add all your Animal Feature bonuses to a single thing - like your nose, or one of your hands or something - then you're clearly acting counter to the spirit of the boon and the Animal purview as a whole by attempting to gain all its benefits while not actually becoming very animal-like. You get those benefits because you're more like an animal, not in spite of it, so trying to weasel out of being critter-like because it's socially smarter or easier to hide that way won't cut it. You're in training to be a god of animals; you don't get to confine all your animal-ness to a tail that you hide in your pants all the time anyway. That's a fundamental failure at the point of the boon, which is to draw upon the beast-like nature of your powers to enhance your performance.

It's also no fun if you're just combining features because you can't think of any other feature to use, or because you're avoiding tarnishing an "image" of yourself that doesn't include creature parts. If your mental image of what your character should be like can't include animal body parts, you probably need to rethink your decision to invest in the Animal purview at all - what are you even doing with it, in that case? If you only took it because you wanted the sweet bonuses, suck it up and start picking some features to claw or fur or scale up. If you took it because you want to be the commander-of-beasts kind of animal god instead of the embodier of their animal nature, that's totally fine, but that boils down to a roleplaying decision; if you want to appear as more human at a particular moment for whatever reason, don't use the boons that make you more animalistic right then. Just like the folks with Sun sometimes don't use their powers because being lit up like a runway can be inconvenient or counter to what they're doing, you may not always be able to conveniently animalize yourself without attracting notice. If you legitimately aren't sure what other part of you would make a good choice, or you have a really hard-to-use animal (like Jellyfish, for example), getting together with the Storyteller or other players can help spark new ideas.

Basically, this is a rule-of-awesomeness moment: if you're doing it because it's awesome, then yes. If you're doing it for any other reason - it's more convenient, it's less noticeable, you don't really like Animal but you do like dice bonuses, or anything else - then no. The Animal purview is about animals, not dice, regardless of how some powers are mechanically represented in the system. If you don't want to be involved in animal stuff, you don't want this purview.

So for your examples, we might allow the eyes, if you had a cool reason why each individual eye was doing something different, but not the nose, which is a clear example of double-dipping with no enhancement from having both "features" in the same place. As a general rule, we encourage players to choose very different features for each bonus unless the animal in question gives them a reason not to, so most of the time eyes, hands, ears and other paired organs or body parts are considered one "feature" together.